The Washington State University Catalog

Courses with the BIOLOGY Subject

The online catalog includes the most recent changes to courses and degree requirements that have been approved by the Faculty Senate, including changes that are not yet effective.

Courses

The online catalog includes the most recent changes to courses and degree requirements that have been approved by the Faculty Senate, including changes that are not yet effective. Courses showing two entries of the same number indicate that the course information is changing. The most recently approved version is shown first, followed by the older version, in gray, with its last-effective term preceding the course title. Courses shown in gray with only one entry of the course number are being discontinued. Course offerings by term can be accessed by clicking on the term links when viewing a specific campus catalog.


Biology (BIOLOGY)

Fall 2020 Spring 2021 


101 [BSCI] Biology of Humans 3 The biology of good health and longevity; evaluation of lifestyle choices; consideration of each body system and the potential for disease and disorder. Credit not granted towards elective requirements for majors in the School of Biological Sciences. Typically offered Fall, Spring, and Summer.

102 [BSCI] General Biology 4 (3-3) Enrollment not allowed if credit for BIOLOGY 105 already earned. Understanding current and future advances in biology as 'citizen scientists'. Lecture and laboratory; not for students majoring in the life sciences. Credit not allowed for students who have already completed BIOLOGY 105. Credit not granted towards elective requirements for majors in the School of Biological Sciences. Typically offered Fall, Spring, and Summer.

103 Science and Scientific Thinking 1 (0-3) Exploring science as a tool for understanding nature using case studies, experimentation, and data analysis. Topics range from atoms to ecosystems including physiology, inheritance, and the carbon cycle. Credit not granted towards elective requirements for majors in the School of Biological Sciences. Recommended for students with an ALEKS math placement score of less than 45%. (Crosslisted course offered as BIOLOGY 103, SCIENCE 103). Typically offered Fall.

105 General Biology Laboratory 1 (0-3) Course Prerequisite: Junior standing. Enrollment not allowed if credit for BIOLOGY 102 already earned. Understanding biology as a science and its effect on issues within society. Laboratory only. Credit not allowed for students who have already completed BIOLOGY 102. Credit not granted towards elective requirements for majors in the School of Biological Sciences. Typically offered Fall, Spring, and Summer.

106 [BSCI] Introductory Biology: Organismal Biology 4 (3-3) Course Prerequisite: One of the following -- a minimum ALEKS math placement score of 40%, MATH 100 with an S, MATH 101 with a C or better, MATH 103 or higher, BIOLOGY 103 with a C or better, BIOLOGY 102, BIOLOGY 120, or 3 credits of biology with a lab. One semester of a two semester sequence (BIOLOGY 106/107 or BIOLOGY 107/106) for science majors and pre-professional students. Biology of organisms; plants, animals, ecology and evolution. Typically offered Fall, Spring, and Summer.

107 [BSCI] Introductory Biology: Cell Biology and Genetics 4 (3-3) Course Prerequisite: Minimum 2 credits 100 level CHEM or concurrent enrollment. First or second semester of a one-year sequence (BIOLOGY 106/107 or BIOLOGY 107/106) for science majors and pre-professional students. Cell biology and genetics of prokaryotes and eukaryotes. Typically offered Fall, Spring, and Summer.

110 Biological Perspectives on Environmental Issues 3 Current case studies of human interaction with the environment exploring concepts in ecology, biodiversity, global chemical cycles, and climate change. Credit not granted towards elective requirements for majors in the School of Biological Sciences.

111 [BSCI] Laboratory Experiments in Biology and Genetics 1 (0-3) Scientific method and its application to a diverse range of biology and genetics topics and research questions. Credit not granted towards elective requirements for majors in the School of Biological Sciences.

120 [BSCI] Introductory Botany 4 (3-3) Introduction to plant science, highlighting certain aspects of plant biology and current research and how these relate to us all in the modern world. Credit not granted towards elective requirements for majors in the School of Biological Sciences. Typically offered Fall.

125 Genetics and Society 3 Genetic topics in media and daily life including human health, agriculture, ecology and forensics for the educated non-biologist. Credit not granted towards elective requirements for majors in the School of Biological Sciences.

135 Animal Natural History 3 Identification, life history, habitat relations, ecology, behavior, and conservation of animals commonly found in the Pacific Northwest. Credit not granted towards elective requirements for majors in the School of Biological Sciences.

140 [BSCI] Introduction to Nutritional Science 3 Information related to dietary sources of nutrients, their functions in the body, physiologic and environmental factors that govern nutrient requirements, and guidelines for optimal dietary patterns. Credit not granted towards elective requirements for majors in the School of Biological Sciences. Typically offered Fall, Spring, and Summer.

150 [BSCI] Evolution 3 Basic principles and implications of Darwinian evolution. Credit not granted towards elective requirements for majors in the School of Biological Sciences.

201 Contemporary Biology 1 Course Prerequisite: BIOLOGY 101, 102, 106, 107, 120, or MBIOS 101. Biological information that provides a framework for understanding life processes; impact of biological information on human affairs. Typically offered Fall and Spring.

210 Your Future in Life Sciences 2 Exploration of career options in biological sciences with faculty and outside speakers; guide to preparing resume and career plans. (Crosslisted course offered as SCIENCE 210, BIOLOGY 210).

220 Medical Terminology 2 Course Prerequisite: BIOLOGY 101, 102, 106, 107, or KINES 262. Terms and word constructions for health care occupations; format and function of medical records. Typically offered Spring.

221 Exploring Health Careers 1 Introduction to human and animal health care careers. S, F grading.

225 Preparation for the Health Care Workplace 2 (1-3) Basic content and skills to prepare for health related internships.

251 Introductory Human Physiology 4 (3-3) Course Prerequisite: BIOLOGY 102, 106, or 107. Basic physiological processes in humans from the cellular to the organismal level. Credit not granted for both BIOLOGY 251 and 353. Typically offered Fall and Summer.

298 [BSCI] Honors Biology for Non-Science Majors 4 (3-3) Course Prerequisite: Must be an Honors student. Understanding the natural world from a biological perspective for non-science majors. Typically offered Spring.

301 General Genetics 4 Course Prerequisite: BIOLOGY 106 or 120; BIOLOGY 107; CHEM 101 or 105; CHEM 102 or 106. Principles of modern and classical genetics. (Crosslisted course offered as MBIOS 301, BIOLOGY 301). Typically offered Fall, Spring, and Summer.

307 [DIVR] Biology of Women 3 Course Prerequisite: BIOLOGY 102 or 106. Biological basis of sex and its relationship to body function, women and health care, and the impact of social and cultural perspectives on the experience of being female. (Crosslisted course offered as BIOLOGY 307, WOMEN ST 307). Typically offered Spring.

308 [BSCI] Marine Biology 3 Course Prerequisite: BIOLOGY 106; sophomore standing. Introduction to the marine environment including oceanic, near-shore and estuarine communities of organisms and their roles and interactions.

315 Gross and Microanatomy 4 (3-3) Course Prerequisite: At least 3 hours of BIOLOGY; sophomore standing; cumulative WSU GPA 2.5 or better. Gross and microscopic anatomy of the human body. Recommended for pre-health care professionals only. Typically offered Fall and Spring.

321 [M] Principles of Animal Development 4 (3-3) Course Prerequisite: BIOLOGY 106; BIOLOGY 107. Experimental analyses of development and descriptive and comparative examination of embryology; emphasis on the chordates. Recommended preparation: BIOLOGY 301 or MBIOS 301. Typically offered Fall.

322 [M] Invertebrate Biology 4 (3-3) Course Prerequisite: BIOLOGY 106. Phylogenetic relationships, development, and functional ecology of the invertebrate animals. Typically offered Even Years - Fall.

324 Comparative Vertebrate Anatomy 4 (2-6) Course Prerequisite: BIOLOGY 106. Evolution of vertebrates and their organ systems; correlation of structural modification with function. Typically offered Spring. Cooperative: Open to UI degree-seeking students.

330 Principles of Conservation 3 Course Prerequisite: BIOLOGY 102, 106, or 107. Conservation of major natural resources through a biological approach; philosophical, economic, and political aspects of important conservation issues. Typically offered Spring and Summer.

332 [M] Systematic Botany 4 (3-3) Course Prerequisite: BIOLOGY 106 or 120. Identification and classification of vascular plants with emphasis on the local flora. Typically offered Odd Years - Spring.

333 [BSCI] Human Nutrition and Health 3 Course Prerequisite: BIOLOGY 102, 106, 107, 251, 315, or concurrent enrollment in BIOLOGY 251. Credit not granted for students who have already completed BIOLOGY 233 with a grade of C or above. Foundations in nutritional science and its relationship to human health through the application of fundamental principles of biology. Typically offered Fall and Spring.

335 [M] Genome Biology 3 Course Prerequisite: BIOLOGY 301. Comparative analysis of genomes from bacteria to humans including methods for sequencing, genotyping, annotation of genomes, population genetics and evolution. Typically offered Fall.

340 Introduction to Mathematical Biology 3 Course Prerequisite: MATH 140 with a C or better, or MATH 172 with a C or better, or MATH 182 with a C or better; BIOLOGY 101, BIOLOGY 102, BIOLOGY 106, or BIOLOGY 107. Mathematical biology and development of mathematical modeling for solutions to problems in the life sciences. (Crosslisted course offered as MATH 340, BIOLOGY 340). Typically offered Spring.

350 Comparative Physiology 4 (3-3) Course Prerequisite: BIOLOGY 107; CHEM 345. Analysis of systems and integrative physiology with an emphasis on evolutionary adaptation among mammalian and non-mammalian vertebrates. Typically offered Fall.

352 Cells 3 Course Prerequisite: BIOLOGY 107; CHEM 345. Diversity and processes at the cellular level; structure and function. Typically offered Fall and Spring.

353 Advanced Human Physiology 4 (3-3) Course Prerequisite: BIOLOGY 106; BIOLOGY 107. Function and control at the organ-organismic level with emphasis on mammals, including humans; emphasis on human health science applications. Credit not granted for both BIOLOGY 251 and 353. Recommended preparation: BIOLOGY 315 or 354. Typically offered Spring.

354 Human Anatomy for Health Occupations 4 (3-3) Course Prerequisite: BIOLOGY 107; CHEM 102 or 345. History and anatomy of humans with non-cadaver-based laboratory utilizing preserved and histological specimens, models and software.

360 Molecular Processes of Living Organisms 3 Course Prerequisite: BIOLOGY 107. Exploration of fundamental molecular processes to encourage thinking beyond biological species in order to comprehend larger-scale biological issues and relevance for society. Typically offered Even Years - Fall.

370 [M] Ecology of Health and Disease 4 (3-3) Course Prerequisite: BIOLOGY 106; CHEM 102 or 105. Enrollment in BIOLOGY 370 not allowed if credit already earned for BIOLOGY 372. Ecology of species interactions in changing environments and how they influence human and animal health. Credit not granted for both BIOLOGY 370 and 372. Field trips may be required.

372 [M] General Ecology 4 (3-3) Course Prerequisite: BIOLOGY 106; CHEM 102 or 105. Enrollment in BIOLOGY 372 not allowed if credit already earned for BIOLOGY 370. Relationship of organisms with physical and biotic components of their environment at the population, community, and ecosystem level. Credit not granted for both BIOLOGY 370 and 372. Field trips may be required. Typically offered Fall, Spring, and Summer.

390 Stream Monitoring 1 (0-3) Course Prerequisite: BIOLOGY 101, 102, or 106; CHEM 101 or 105; junior standing. Principles and methods of water quality monitoring, including habitat assessment, water chemistry, and biological assessment. Field work and independent research required.

393 [M] Professional Communications in Biology 2 Course Prerequisite: Admitted to the major in Biology or Zoology. Literature investigation, oral presentation and written reports of selected topics in biology. Typically offered Fall and Spring.

394 Medicine as a Career 2 Course Prerequisite: Junior standing. Current issues in medicine; ethical, financial, and personal aspects of medical practice. Typically offered Spring. S, F grading.

395 Evolutionary Medicine 3 Course Prerequisite: BIOLOGY 301. Modern medical issues from an evolutionary perspective, integrated with other biological fields in medical research; topics include disease diversity, immune function, the evolution of virulence, human disease management, cancer, obesity, and human mental and reproductive health issues and their management. Typically offered Odd Years - Spring.

401 [CAPS] Plants and People 3 Course Prerequisite: BIOLOGY 106 or 120; BIOLOGY 107; junior standing. Relationships between plants and people, especially cultural and economic applications of plants. Typically offered Spring.

402 [M] Beneficial Microbes in Nature and Society 3 Course Prerequisite: BIOLOGY 372, 403, or 405; junior standing. In-depth investigations of interdisciplinary topics addressing the importance of beneficial microbes to organisms, natural systems, and society from across the disciplines of microbiology, medicine, evolutionary ecology, and agricultural science. Typically offered Odd Years - Spring.

403 Evolutionary Biology 3 Course Prerequisite: BIOLOGY 301. The survey of evidence for evolution and operation of evolutionary processes that influence adaptation, diversification and speciation in organisms.

405 Principles of Organic Evolution 3 (2-3) Course Prerequisite: BIOLOGY 301. The evolutionary processes that influence adaptation, population differentiation, and speciation in organisms. Typically offered Spring.

408 [CAPS] [M] Contemporary Genetics 3 Course Prerequisite: MBIOS / BIOLOGY 301 with a C or better; junior standing. Consideration of the state-of-the-art genetic technologies and their impact on society, environment and the economy. Typically offered Spring.

409 Plant Anatomy 4 (2-6) Course Prerequisite: BIOLOGY 106 or 120. Developmental anatomy and morphology of vascular plants; economic forms. Credit not granted for both BIOLOGY 409 and BIOLOGY 509. Offered at 400 and 500 level. Typically offered Even Years - Fall.

410 Marine Ecology 3 Course Prerequisite: BIOLOGY 106. The ecology and conservation of marine organisms, communities, and ecosystems. Typically offered Fall.

412 Biology of Fishes 3 (2-3) Course Prerequisite: BIOLOGY 106. Evolution, identification, life history, and characteristics of important fish species.

418 Parasitology 4 (3-3) Course Prerequisite: BIOLOGY 102 or BIOLOGY 106; junior standing. Types of associations, life cycles, control, prevention, and modifications of parasites; examination of parasitic protozoa and helminths. Typically offered Fall.

420 Plant Physiology 3 Course Prerequisite: BIOLOGY 106 or 120. Water relations, mineral nutrition, photosynthesis, respiration, and growth of plants. Recommended: Organic chemistry. Typically offered Fall.

421 Plant Physiology Laboratory 1 (0-3) Course Prerequisite: BIOLOGY 420 or concurrent enrollment. Laboratory for Biol 420.

423 Ornithology 4 (3-3) Course Prerequisite: BIOLOGY 106. Ecology, systematics, and evolution of birds. Field trips required include two Saturdays. Typically offered Spring.

428 Mammalogy 4 (3-3) Course Prerequisite: BIOLOGY 106. Ecology, systematics, and evolution of mammals. Typically offered Fall.

430 Methods of Teaching Secondary Science I 3 Course Prerequisite: Junior standing. Application of learning and theory and philosophy and structure of science in teaching middle and secondary school science courses. (Crosslisted course offered as BIOLOGY 430, MBIOS 480, TCH LRN 430). Typically offered Fall.

431 Methods of Teaching Secondary Science II 3 Course Prerequisite: BIOLOGY 430, MBIOS 480, or TCH LRN 430; junior standing. Integration of assessment, curricular, and technological tools into instruction that aligns with learning theory and the philosophy/structure of science. (Crosslisted course offered as BIOLOGY 431, MBIOS 481, TCH LRN 431). Typically offered Spring.

432 [M] Biology of Amphibians and Reptiles 4 (3-3) Course Prerequisite: BIOLOGY 106; BIOLOGY 372 or SOE 300. Characteristics, evolution, and systematics; patterns of distribution; adaptive strategies; interactions between humans and amphibians and reptiles. Typically offered Even Years - Spring.

438 [M] Animal Behavior 3 (2-3) Course Prerequisite: BIOLOGY 106. Biological study of animal behavior as viewed from ethological, genetic, developmental, ecological, and evolutionary perspectives. Typically offered Fall and Spring.

446 Mutualism and Symbiosis 3 Course Prerequisite: BIOLOGY 372, 403, or 405. Critical evaluation of the ecology, evolution, and molecular biology of mutualism and symbiosis. Credit not granted for both BIOLOGY 446 and 546. Offered at 400 and 500 level. Typically offered Even Years - Fall and Summer.

456 Neuroethology 3 Course Prerequisite: BIOLOGY 301, MBIOS 303, or 300-level NEUROSCI course; STAT 412 or concurrent enrollment. Introduction to neural mechanisms underlying natural animal behaviors from the cellular level to the organismal level.

462 Community Ecology 3 Course Prerequisite: BIOLOGY 372 with a C or better. Assembly, essential properties, levels of interactions, succession, and stability of natural communities; emphasizes an experimental approach to community investigation. Credit not granted for both BIOLOGY 462 and BIOLOGY 562. Recommended preparation: BIOLOGY 372. Offered at 400 and 500 level. Typically offered Fall.

465 Field Stream Ecology 2 Course Prerequisite: BIOLOGY 372. Ecological roles of immature insects in different size streams; pattern changes along the stream continuum; other ecological characteristics.

469 [M] Ecosystem Ecology and Global Change 3 Historic and current factors controlling the function of ecosystems and their responses to natural and human caused global change. Credit not granted for both BIOLOGY 469 and 569. Offered at 400 and 500 level. Typically offered Odd Years - Spring. Cooperative: Open to UI degree-seeking students.

470 Diversity of Plants 3 Morphological, life history, and ecological diversity of major plant clades; emphasis on principles of homology, character transformation, and macroevolution.

473 [CAPS] [M] Evolution and Society 3 Course Prerequisite: ANTH 260 or BIOLOGY 301; junior standing. Survey of how the theory of evolution is used to better understand ourselves, the societies in which live, and the biological world on which we depend. Recommended preparation: BIOLOGY 405 or concurrent enrollment. (Crosslisted course offered as BIOLOGY 473, ANTH 473).

474 Computational Biology 4 (3-3) Course Prerequisite: BIOLOGY 301; MATH 140 or 171; STAT 212, 412, or PSYCH 311. Theory and current literature on a wide range of computational techniques used to address and solve problems in biology; a practical introduction to R/python as scientific languages useful in the solution of problems in biology. Typically offered Odd Years - Spring.

475 Systems Biology of Reproduction 3 Current literature based course on systems biology with a molecular/epigenetic to physiological level understanding of cell, development, disease, and evolutionary biology. Credit not granted for both BIOLOGY 475 and 575. Offered at 400 and 500 level. Typically offered Even Years - Spring.

476 Epigenetics and Systems Biology 3 Course Prerequisite: BIOLOGY 301. Current literature based course on epigenetics and systems biology with topics in environmental epigenetics, disease etiology, and role epigenetics in evolutionary biology. Credit not granted for both BIOLOGY 476 and 576. Offered at 400 and 500 level. Typically offered Odd Years - Spring.

480 [M] Writing in Biology 2 Course Prerequisite: Admitted to the major in Biology or Zoology. Discussion and practice in relating thinking and writing; popular and professional communication in biology.

483 [CAPS] [M] Organisms and Global Change 3 Course Prerequisite: BIOLOGY 372; junior standing. Interaction between organisms and global change across scales of biology. Typically offered Fall.

485 [CAPS] Biology of the Oceans 3 Course Prerequisite: BIOLOGY 106; junior standing. Interdisciplinary capstone course that explores the ocean world from molecules to ecosystems in the context of scientific discovery and society.

486 [M] Marine Invertebrate Communities 3 (2-3) Course Prerequisite: BIOLOGY 106. Survey of marine invertebrates and their habitats. One-week field/lab course at a marine station. Typically offered Summer Session.

489 [CAPS] [M] Synthesis and Communication of Independent Research 3 Course Prerequisite: 2 credits BIOLOGY 499; admitted to major in Biology or Zoology; junior standing; by permission only. Integration of broad topics from biology and other science fields to inform scientific writing and presentation of independent research projects.

490 [M] Professional Seminar in Physical Therapy 2 Course Prerequisite: By permission only. Consideration of treatment modalities and health issues in physical therapy and related disciplines. Typically offered Fall and Spring. A, S, F grading.

491 Clinical Experience V 1-4 May be repeated for credit; cumulative maximum 20 hours. Course Prerequisite: PSYCH 105; BIOLOGY 315; junior standing; by permission only. Work experience under supervision of a qualified professional in a clinical setting. Typically offered Fall, Spring, and Summer. S, F grading.

492 Topics in Biology V 1-3 May be repeated for credit; cumulative maximum 6 hours.

494 Seminar in Mathematical Biology 1 May be repeated for credit; cumulative maximum 4 hours. Course Prerequisite: MATH 140 with a C or better, or MATH 172 with a C or better, or MATH 182 with a C or better; BIOLOGY 101, BIOLOGY 102, BIOLOGY 106, or BIOLOGY 107. Oral presentation of research approaches, research results and literature review of mathematical biology including mathematical modeling of biological systems. (Crosslisted course offered as MATH 494, BIOLOGY 494). Typically offered Spring. Cooperative: Open to UI degree-seeking students. S, F grading.

495 Internship in Biology, Botany, and Zoology V 1-4 May be repeated for credit; cumulative maximum 8 hours. Course Prerequisite: By permission only. Experience in work related to specific career interests. Typically offered Fall, Spring, and Summer. S, F grading.

496 [M] Special Problems and Reports V 1-4 Course Prerequisite: By permission only. Independent project with written project proposal, progress report, and final report required. Typically offered Fall, Spring, and Summer. S, F grading.

497 Instructional Practicum V 1-4 May be repeated for credit; cumulative maximum 8 hours. Academic traineeship in laboratory teaching and tutoring. Typically offered Fall and Spring.

499 Special Problems V 1-4 May be repeated for credit. Course Prerequisite: By permission only. Independent study conducted under the jurisdiction of an approving faculty member; may include independent research studies in technical or specialized problems; selection and analysis of specified readings; development of a creative project; or field experiences. Typically offered Fall, Spring, and Summer. S, F grading.

500 Seminar 1 May be repeated for credit. Typically offered Fall, Spring, and Summer. S, F grading.

501 Proposal Defense Seminar 2 Research proposal defense as part of the preliminary examination for candidacy in the Ph.D. program. Typically offered Fall, Spring, and Summer. S, F grading.

504 Experimental Methods in Plant Physiology 4 (2-6) Advanced techniques and instrumental methods applicable to research in plant physiology. Typically offered Odd Years - Fall.

509 Plant Anatomy 4 (2-6) Developmental anatomy and morphology of vascular plants; economic forms. Credit not granted for both BIOLOGY 409 and BIOLOGY 509. Offered at 400 and 500 level. Typically offered Even Years - Fall.

512 Molecular Mechanisms of Plant Development 3 Physiology of growth; metabolism during development and reproduction.

513 Plant Metabolism 3 Metabolic processes unique to plants, including the primary incorporation of nitrogen, sulfur, carbon dioxide and phosphate into bio-molecules. Typically offered Even Years - Fall.

514 Fish Genetics 2 Chromosomal, biochemical, quantitative, and ecological aspects of fish genetics with emphasis on applications to aquaculture and fish management. Cooperative: Open to UI degree-seeking students.

517 Stress Physiology of Plants 3 Temperature, light, salinity, water effects on physiological processes; mechanistic understanding of stress. Typically offered Even Years - Spring.

519 Introduction to Population Genetics 3 Survey of basic population and quantitative genetics. Typically offered Fall. Cooperative: Open to UI degree-seeking students.

520 Conservation Genetics 2 Genetic studies and approaches relevant to efforts to conserve threatened and endangered populations of organisms.

521 Quantitative Genetics 3 Course Prerequisite: BIOLOGY 519. Fundamentals of quantitative genetics; evolutionary quantitative genetics. Cooperative: Open to UI degree-seeking students.

531 Principles of Systematic Biology 3 Systematic theory; history and current views; approaches to phylogenetic analysis and classification.

533 Modern Methods in Phylogenetics 4 (2-6) Selecting, gathering, and analyzing morphological, cytological, molecular data for phylogenetic and evolutionary studies. Typically offered Even Years - Fall.

534 Modern Methods in Population Genomics 3 Course Prerequisite: BIOLOGY 519. Problems and prospects of designing a study with genomic data: from raw data to demography and selection inferences. Typically offered Odd Years - Spring.

537 Plant Cell Biology 3 Structure and function of plant cells including membrane biology, protein targeting and molecular signaling with emphasis on current research. Typically offered Odd Years - Spring.

540 Stable Isotope Theory and Methods 3 Theory and practice of measuring stable isotope ratios of biologically important elements. Typically offered Odd Years - Fall. Cooperative: Open to UI degree-seeking students.

544 Nitrogen Cycling in the Earth's Systems 3 Nitrogen dynamics in terrestrial, aquatic, and atmospheric systems; nitrogen transformations in natural and managed systems and responses to human activities. (Crosslisted course offered as BIOLOGY 544, SOIL SCI 544).

545 Statistical Genomics 3 (2-3) Develop concepts and analytical skills for modern breeding by using Genome-Wide Association Study and genomic prediction in framework of mixed linear models and Bayesian approaches. (Crosslisted course offered as CROP SCI 545, ANIM SCI 545, BIOLOGY 545, HORT 545, PL P 545.) Recommended preparation: BIOLOGY 474; MBIOS 478. Typically offered Spring. Cooperative: Open to UI degree-seeking students.

546 Mutualism and Symbiosis 3 Critical evaluation of the ecology, evolution, and molecular biology of mutualism and symbiosis. Credit not granted for both BIOLOGY 446 and 546. Offered at 400 and 500 level. Typically offered Even Years - Spring.

548 Evolutionary Ecology of Populations 3 Evolutionary dynamics of natural populations and the co-evolution of species. Typically offered Odd Years - Spring. Cooperative: Open to UI degree-seeking students.

549 Behavioral Ecology 3 Examination of animal behavior from evolutionary and ecological perspectives. Typically offered Even Years - Fall.

556 Biochemical Adaptation 3 Relationships between enzyme/macromolecule adaptation and animal performance.

559 Hormones, Brain and Behavior 3 Classical behavioral endocrinology from molecular to whole organisms, integrating evolutionary ecology, neuroethology and behavioral neuroendocrinology. Typically offered Even Years - Fall.

560 Plant Ecophysiology 3 Relationships of biotic and abiotic environment to plant distribution and evolution through study of physiological processes.

561 Environmental Physiology 3 Individual and evolutionary adaptations to changing environments with emphasis on recent literature. Typically offered Even Years - Spring.

562 Community Ecology 3 Assembly, essential properties, levels of interactions, succession, and stability of natural communities; emphasizes an experimental approach to community investigation. Credit not granted for both BIOLOGY 462 and BIOLOGY 562. Recommended preparation: BIOLOGY 372. Offered at 400 and 500 level. Typically offered Fall.

563 Field Ecology 2 (0-6) Field implementation of descriptive and experimental techniques to quantify the structure, composition, and interactions within natural communities. Field trips required. Typically offered Even Years - Spring. Cooperative: Open to UI degree-seeking students.

564 Molecular Ecology and Phylogeography 3 Use of genetic markers for the study of ecological phenomena, including kinship, population structure, and phylogeography. Typically offered Even Years - Fall. Cooperative: Open to UI degree-seeking students.

565 Ecology and Evolution of Disease 3 Disease ecology and evolution with a focus on current literature. Recommended preparation: BIOLOGY 372; BIOLOGY 405. Typically offered Odd Years - Fall. Cooperative: Open to UI degree-seeking students.

566 Mathematical Genetics 3 Mathematical approaches to population genetics and genome analysis; theories and statistical analyses of genetic parameters. (Crosslisted course offered as MATH 563, BIOLOGY 566). Required preparation must include multivariate calculus, genetics, and statistics. Typically offered Odd Years - Fall. Cooperative: Open to UI degree-seeking students.

567 Ecological Restoration 3 Introduction to major issues in restoration ecology; major ecological dimensions of restoration.

568 Conservation Ecology 3 Diagnosis of endangered species, population viability analysis, invasive species ecology, landscape ecology and ecosystem management.

569 [M] Ecosystem Ecology and Global Change 3 Historic and current factors controlling the function of ecosystems and their responses to natural and human caused global change. Credit not granted for both BIOLOGY 469 and 569. Offered at 400 and 500 level. Typically offered Odd Years - Spring. Cooperative: Open to UI degree-seeking students.

570 Diversity of Plants 3 Morphological, life history, and ecological diversity of major plant clades; emphasis on principles of homology, character transformation, and macroevolution.

571 Quantitative Toolkit for Biologists 3 Course Prerequisite: STAT 512. Hands-on experience in the exploration, analysis, and interpretation of patterns in modern biological datasets. Typically offered Even Years - Spring.

572 Quantitative Methods and Statistics in Ecology 4 (3-3) Course Prerequisite: By permission only. Philosophy and methods of formulating hypotheses as mathematical models and confronting them with data. Typically offered Odd Years - Spring.

573 Ancient DNA 3 The prospects and problems associated with the study of ancient DNA are explored through reading and discussing primary literature. Typically offered Even Years - Spring.

575 Systems Biology of Reproduction 3 Current literature based course on systems biology with a molecular/epigenetic to physiological level understanding of cell, development, disease, and evolutionary biology. Credit not granted for both BIOLOGY 475 and 575. Offered at 400 and 500 level. Typically offered Even Years - Spring.

576 Epigenetics and Systems Biology 3 Current literature based course on epigenetics and systems biology with topics in environmental epigenetics, disease etiology, and role epigenetics in evolutionary biology. Credit not granted for both BIOLOGY 476 and 576. Offered at 400 and 500 level. Typically offered Odd Years - Spring.

579 Mathematical Modeling in the Biological and Health Sciences 3 Techniques, theory, and current literature in mathematical modeling in the biological and health sciences, including computational simulation. (Course offered as BIOLOGY 579, MATH 579). Typically offered Odd Years - Fall. Cooperative: Open to UI degree-seeking students.

581 Comparative Biology of Social Traditions 3 Phylogenetic and modeling perspectives used to examine the evolution of social learning and cultural transmission in humans and other animals. (Crosslisted course offered as ANTH 581, BIOLOGY 581).

582 Professional Communication in Biology - Grant Writing 2 Mechanics and style of publishing biological research and findings; adaptation of writing to various venues and audiences with emphasis on grant writing. Typically offered Spring.

585 Professional Development and Training for College and University Teaching 2 Preparation for roles as teaching assistants and as instructors of undergraduate classroom education. Typically offered Fall.

589 Advanced Topics in Biology V 1-3 May be repeated for credit; cumulative maximum 6 hours. Recent advances in biology.

591 Seminar in Molecular Plant Sciences 1 May be repeated for credit; cumulative maximum 4 hours. A cross-discipline seminar, including botany, crop and soils sciences, horticulture, plant pathology, and molecular plant sciences.

593 Seminar I 1 May be repeated for credit. Literature and problems. Typically offered Fall and Spring.

597 Teaching Practicum V 1-4 May be repeated for credit; cumulative maximum 4 hours. Zoology laboratory teaching internship. S, F grading.

600 Special Projects or Independent Study V 1-18 May be repeated for credit. Independent study, special projects, and/or internships. Students must have graduate degree-seeking status and should check with their major advisor before enrolling in 600 credit, which cannot be used toward the core graded credits required for a graduate degree. S, F grading.

700 Master's Research, Thesis, and/or Examination V 1-18 May be repeated for credit. Independent research and advanced study for students working on their master's research, thesis and/or final examination. Students must have graduate degree-seeking status and should check with their major advisor/committee chair before enrolling for 700 credit. Typically offered Fall, Spring, and Summer. S, U grading.

702 Master's Special Problems, Directed Study and/or Examination V 1-18 May be repeated for credit. Independent research in special problems, directed study, and/or examination credit for students in a non-thesis master's degree program. Students must have graduate degree-seeking status and should check with their major advisor/committee chair before enrolling for 702 credit. S, U grading.

800 Doctoral Research, Dissertation, and/or Examination V 1-18 May be repeated for credit. Course Prerequisite: Admitted to the Biology, Plant Biology, Botany, or Zoology PhD program. Independent research and advanced study for students working on their doctoral research, dissertation and/or final examination. Students must have graduate degree-seeking status and should check with their major advisor/committee chair before enrolling for 800 credit. Typically offered Fall, Spring, and Summer. S, U grading.

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