The Washington State University Catalog

Program in Neuroscience

The online catalog includes the most recent changes to courses and degree requirements that have been approved by the Faculty Senate, including changes that are not yet effective.

Program in Neuroscience

ipn.vetmed.wsu.edu/neuroscience/
Veterinary and Biomedical Research Bldg (VBR) 207
509-335-6624

Director, Chair, and Professor, S. Hentges; Associate Director of Graduate Program and Professor, S. Appleyard; Associate Director of Undergraduate Program and Associate Professor, S. Gizerian; Regents Professor, J. M. Krueger; Professors, G. Belenky (Spokane), M. Chandra, R. M. Craft, M. Frank (Spokane), R. Fuchs, J. Hinson, M. Morgan (Vancouver), C. Portfors (Vancouver), R. Quock, R. C. Ritter, M. Schmitter-Edgecombe, H. Van Dongen (Spokane), G. Wayman, P. Whitney; Associate Professors, R. L. Brown, T. Brown, A. Coffin (Vancouver), C. Davis (Spokane), A. Dimitrov (Vancouver), W. Dong, D. Jackson (Spokane), L. Kapas (Spokane), M. Layton (Spokane), D. Lin, R. McLaughlin, S. McPherson (Spokane) P. Meighan, J. Peters, D. Rossi, L. Sprunger, E. Szentirmai (Spokane), B. Tanner, M. Varnum, A. Vasavada, J. Wisor (Spokane); Assistant Professors, R. Catena, K. Delevich, J. Gerstner (Spokane), A. Henricks, K. Honn, L. Peixoto (Spokane), E. Qualls-Creekmore, B. Satterfield (Spokane); Additional Graduate Faculty, C. Burgoyne, S. Demirel, B. Fortune, H. Shen, B. Sorg, D. Stenkamp, L. Wang.

Neuroscience is the study of how the nervous system impacts physiology, behavior, and health. It seeks to answer questions at the molecular, cellular, systems, behavioral, cognitive, and societal levels through application of a wide array of approaches and techniques. Neuroscience research is critical not only to improving learning and performance, but also addresses some of the most vexing problems that prevent good health, as well as impacts the very nature on how we view ourselves as sentient beings. WSU has a neuroscience faculty engaged in numerous focuses with particular strengths in the areas of energy homeostasis, sleep, circadian biology, addiction, emotions and well-being, sensory processes, neuronal function, electrophysiology, and movement. The Program sponsors Bachelor of Science, Master of Science, and Doctor of Philosophy degrees, in addition to minors at the undergraduate level.

The undergraduate program for majors is designed for students interested in preparing for professional study in the health sciences (such as medical doctor or doctor of veterinary medicine), graduate school, or for those who wish to use their training in laboratory settings in universities, government organizations, or industry.

Computational neuroscience is an option within the undergraduate neuroscience major and links the information processing features of the nervous system with information processing of computer systems. Accordingly, the computational neuroscience track supplements the neuroscience core curriculum with information technology courses. In this way students learn not only of the brain and its information processing mechanisms, but also of modern computer hardware and software technologies. Upon completion of the four-year curriculum, a BS in Neuroscience will be awarded. Furthermore, the program is designed to allow students to acquire breadth in computation subjects or, alternatively, to focus on either software or hardware aspects of computation. Students choosing to acquire breadth in computational subjects will be well prepared for graduate study in most areas of neural and biomedical science, including bioengineering. Students choosing a software or hardware focus may obtain a minor in either computer science or computer engineering.

Student Learning Outcomes for BS neuroscience majors

Knowledge:

  • Demonstrate knowledge of, and recognize the relationships between, the structure and function of molecules and tissues involved in neurobiological systems at all levels: molecular, cellular, and organismal.
  • Recognize the impact that science has on culture, and vice versa.

Skills:

  • Perform basic laboratory techniques used in neuroscience research and understand and apply principles of laboratory safety.
  • Locate and retrieve scientific information and read, understand, and critically evaluate primary literature.
  • Prepare oral and written reports in a standard scientific format.
  • Apply the scientific process, including designing, conducting, and evaluating experiments and testing of hypotheses.
  • Use mathematics and statistics to evaluate scientific evidence and interpret graphs and tables.

Attitudes

  • Recognize that all areas of science are integrated and interconnected.
  • Appreciate scientific knowledge as something that is not static, but constantly expanding through the ongoing work of researchers.
  • Value ethical conduct in science.
  • Recognize that the best decision-making and policies are based on evidence.

Transfer Students

Transfer students must satisfy the program requirements for graduation. Science courses taken at other institutions will be evaluated and credits accepted where possible. Transfer students are strongly encouraged to enroll in math and chemistry classes at the appropriate level to meet the program requirements prior to transferring. Inquiries should be directed to the Associate Director of Undergraduate Program.

7-Year Honors Neuroscience/Veterinary Medicine Degree Program

Academically qualified undergraduate students who meet the highly selective criteria for admission to WSU’s Veterinary Medical Program may apply to the 7-year BS/DVM degree program in neuroscience after completion of one year of Honors College coursework at WSU. If accepted into the program, the student will work toward a bachelor of science in neuroscience in the first three years of the program and work toward the doctor of veterinary medicine degree in the following four years. The first three years are a combination of WSU Honors College courses and regular university undergraduate courses that fulfill the pre-veterinary and Neuroscience major requirements. The last four years are the traditional doctor of veterinary medicine program, plus completion of an honors thesis. Prospective applicants must be admitted to the WSU Honors College and enrolled in Honors courses. See the Honors College for additional information.

Preparation for Graduate Study in Neuroscience

The graduate program prepares students for careers in academia, industry, and government service. Students work closely with faculty members who are internationally known for their research accomplishments in energy homeostasis, sleep, circadian biology, addiction, emotions and well-being, sensory processes, neuronal function, electrophysiology, and movement.  Upon graduation, students with a doctoral degree are credible international experts in the area of their thesis research. They will have developed cutting edge technical research skills, be capable of independently organizing and writing publishable research manuscripts, know the fundamentals on how to write competitive research proposals, have a breadth of knowledge that enables them to critically evaluate neuroscience research, and finally, develop insights that help them identify areas ripe for future investigation. These skills are not only appropriate for basic research, but the ability to organize a set of sophisticated goals, identify a plan to accomplish those goals, and then successfully complete the plan are skills transferable to many jobs. Our graduates have gone on to excellent positions in tier-one research universities, teaching colleges, industry (e.g., biotechnology, pharmaceutical, and medical device), as well as professional schools (e.g., medical, veterinary, optometry).

To be eligible for admission, candidates must meet general Washington State University requirements outlined at the Graduate School website: http://gradschool.wsu.edu/ in effect at the time of their admission, as well as the current graduate neuroscience program requirements.

Applicants to the Ph.D. or M.S. program are required to have a Bachelor’s degree from an accredited higher education institution. Applicants must have a minimum grade point average of 3.0 (A=4.0).  Applicants will have completed courses in inorganic and organic chemistry, biochemistry, calculus, physics and a minimum of three courses in different areas of the biological sciences. It is advisable that applicants have a basic statistics course prior to entering the neuroscience program.

Application documents must include the following:

  • College transcripts (unofficial acceptable for initial review–upon admittance official transcripts are required)
  • Three (3) letters of reference
  • Resume or curriculum vita
  • Personal statement describing why you are interested in studying neuroscience (clearly define which faculty mentor (minimum of 3) you are interested in working with and explain why). If admitted to WSU you will have the ability to refine your choice of faculty mentors while doing lab rotations.  Included in the personal statement describe an achievement that you are proud of and discuss how you reached your goal, and any obstacles you had to overcome to reach it. Conversely, tell us about a time when you didn’t achieve a goal and what you learned from the experience (maximum word length is 350 words).
  • Writing Statement:  Describe a major finding in neuroscience and/or biomedicine over the past five (5) years, and explain why you think it was important (maximum word length is 350 words for statement).  Be sure to cite references used after the writing statement.  An additional maximum length of 350 words is given for the cite references.
  • Turning in a document over the maximum word length may cause your application to be disqualified from consideration.
  • TOEFL scores (minimum score 100) required for applicants whose native language is not English

Inquiries should be directed to the Program in Neuroscience, Department of Integrative Physiology and Neuroscience; Washington State University, Pullman, WA 99164-7620 or email grad.neuro@wsu.edu

Students normally begin their studies in the fall semester, which starts the latter part of August. Applicants are offered admission on a rolling basis, but may be notified of acceptance as late as April 15. Students may still apply for admission after December, but graduate stipends may not be available for late applicants.




Schedules of Studies

Honors students complete the Honors College requirements which replace the UCORE requirements.


Biomedical Business Option (120 Credits)

Students may be admitted to the Neuroscience - Biomedical Business Option upon making their intention known to the department and enrollment in or credit for MATH 106 (or higher). To maintain admission to the major, students who have completed a minimum of 30 credits at WSU must maintain a 3.0 minimum GPA overall, and a 3.0 minimum GPA in BIOLOGY 107, CHEM 105, CHEM 106 or 116, MATH 140 or 171, NEUROSCI 301, NEUROSCI 302, PHYSICS 101/111 or 201/211 or 205, and PHYSICS 102/112 or 202/212 or 206 or CHEM 345. Students must receive a grade of C or better in these courses and may repeat a maximum of three of these courses to maintain admission to the major.
First Year
First TermCredits
CHEM 105 [PSCI]4
ENGLISH 101 [WRTG]3
MATH 140 [QUAN] or 171 [QUAN]4
NEUROSCI 105 [BSCI]3
PSYCH 105 [SSCI]3
Second TermCredits
BIOLOGY 107 4
CHEM 1064
Communication [COMM] or Written Communication [WRTG]13
HISTORY 105 [ROOT]3
Second Year
First TermCredits
BIOLOGY 1064
CHEM 3454
NEUROSCI 301 23
PHYSICS 101 or 2013
PHYSICS 111 or 2111
Second TermCredits
Arts [ARTS]3
Diversity [DIVR]3
NEUROSCI 3023
PHYSICS 102 or 2023
PHYSICS 112 or 2121
Neuroscience Elective 33
Complete Writing Portfolio
Third Year
First TermCredits
BIOLOGY/MBIOS 3014
Humanities [HUM]3
Behavior Course 43
Statistics Course 53 or 4
Second TermCredits
ENTRP 4903
MBIOS 3034
NEUROSCI 4044
Biomedical Business Elective 63
Fourth Year
First TermCredits
ENTRP 4263
ENTRP 4853
NEUROSCI 430 [M]4
Electives76
Second TermCredits
ENTRP 4863
NEUROSCI 403 [M]3
NEUROSCI 490 [CAPS]3
Electives76

Footnotes
1NEUROSCI/MBIOS 201 is recommended.
2PSYCH 372 may be taken in place of NEUROSCI 301 with department permission.
3Neuroscience Electives (minimum 3 credits): Approved courses include BIOLOGY 307, 315, 321, 333, 340, 352, 353, 354, 438, 456; MATH 340; MBIOS 304, 305, 401, 404, 413; NEUROSCI 305, 333, 409, 425, 426; PHYSICS 466; PSYCH 265, 312, 333, 350, 361, 372, 384, 464, 470, 490, 491; VET PH 308. Other courses may be allowed by department consent. Courses may not be used to fulfill more than one requirement. Please see your advisor.
4Choose one course from: NEUROSCI 305, 333, or 409. Other courses may be allowed by department consent.
5Choose one course from: PSYCH 311, STAT 212, 360, 370, or 412.
6Biomedical Business Electives (minimum 3 credits): Approved courses include ENTRP 489, MGMT 301, MGMT 401, MGMT 483. Other courses may be allowed by department consent. Please see your advisor.
7Elective choices should include 300-400 level coursework to meet the University minimum requirement of 40 upper-division credits.

Computational Breadth of Field Option (124 Credits)

Students may be admitted to the Neuroscience - Computational Breadth of Field Option upon making their intention known to the department and enrollment in or credit for MATH 106 (or higher). To maintain admission to the major, students who have completed a minimum of 30 credits at WSU must maintain a 3.0 minimum GPA overall, and a minimum 3.0 cumulative GPA in BIOLOGY 107, CHEM 105, CHEM 106 or 116, MATH 171, MATH 172, NEUROSCI 301, NEUROSCI 302, and PHYSICS 201/211 or 205. Students must receive a grade of C or better in these courses and may repeat a maximum of three of these courses to maintain admission to the major.
First Year
First TermCredits
Arts [ARTS]3
CHEM 105 [PSCI]4
ENGLISH 101 [WRTG]3
HISTORY 105 [ROOT]3
MATH 171 [QUAN]4
Second TermCredits
BIOLOGY 107 [BSCI] 4
CHEM 1064
CPT S 1214
MATH 1724
Second Year
First TermCredits
BIOLOGY 106 4
MATH 2732
NEUROSCI 30113
PHYSICS 2013
PHYSICS 2111
PSYCH 105 [SSCI]3
Second TermCredits
CHEM 3454
CPT S 1224
Humanities [HUM]3
NEUROSCI 3023
PHYSICS 202 3
PHYSICS 2121
Complete Writing Portfolio
Third Year
First TermCredits
Diversity [DIVR]3
E E 214 4
MATH 2163
MATH 2202
MBIOS / BIOLOGY 3014
Second TermCredits
Communication [COMM] or Written Communication [WRTG]23
MATH 3153
NEUROSCI 4044
Computational Neuroscience Electives33
Fourth Year
First TermCredits
E E 2613
E E 2621
NEUROSCI 4253
NEUROSCI 4261
NEUROSCI 430 [M]4
Computational Neuroscience Electives 33
Second TermCredits
BIO ENG 3404
NEUROSCI 403 [M]3
NEUROSCI 490 [CAPS]3
Computational Neuroscience Electives 33

Footnotes
1PSYCH 372 may be taken in place of NEUROSCI 301 with department permission.
2Professional or Technical Writing is recommended (e.g. ENGLISH 201 or 402)
3Computational Neuroscience Electives (the Breadth of Field emphasis requires a minimum of 9 credits): At least 3 credits must be 300-400-level E E or CPT S courses. Approved courses include BIOLOGY 315, 321, 340, 438, 456; BIO ENG 481; CPT S 322, 434, 440, 443; E E 311, 321, 324, 341, 451, 464; MBIOS 303, 305, 401, 404, 413, 478; NEUROSCI 305, 333, 409; Psych 470, 490, 491; PHYSICS 466. Courses may not be used to fulfill more than one requirement. Other courses may be allowed by department consent. Please consult your advisor.

Computational Hardware Option (124 Credits)

Students may be admitted to the Neuroscience - Computational Hardware Option upon making their intention known to the department and enrollment in or credit for MATH 106 (or higher). To maintain admission to the major, students who have completed a minimum of 30 credits at WSU must maintain a 3.0 minimum GPA overall, and a minimum 3.0 cumulative GPA in BIOLOGY 107, CHEM 105, CHEM 106 or 116, MATH 171, MATH 172, NEUROSCI 301, NEUROSCI 302, and PHYSICS 201/211 or 205. Students must receive a grade of C or better in these courses and may repeat a maximum of three of these courses to maintain admission to the major.
First Year
First TermCredits
Arts [ARTS]3
CHEM 105 [PSCI]4
ENGLISH 101 [WRTG]3
HISTORY 105 [ROOT]3
MATH 171 [QUAN]4
Second TermCredits
BIOLOGY 107 [BSCI]4
CHEM 1064
CPT S 1214
MATH 1724
Second Year
First TermCredits
BIOLOGY 106 4
MATH 2732
NEUROSCI 30113
PHYSICS 2013
PHYSICS 2111
PSYCH 105 [SSCI]3
Second TermCredits
CHEM 3454
CPT S 1224
Humanities [HUM]3
NEUROSCI 3023
PHYSICS 202 3
PHYSICS 2121
Complete Writing Portfolio
Third Year
First TermCredits
BIOLOGY / MBIOS 3014
Communication [COMM] or Written Communication [WRTG]23
E E 214 4
MATH 2163
MATH 2202
Second TermCredits
Diversity [DIVR]3
MATH 3153
NEUROSCI 4044
Computational Neuroscience Electives 33
Fourth Year
First TermCredits
E E 2344
E E 2613
E E 2621
NEUROSCI 4253
NEUROSCI 4261
NEUROSCI 430 [M]4
Second TermCredits
E E 324 [M]4
NEUROSCI 403 [M]3
NEUROSCI 490 [CAPS]3
Computational Neuroscience Electives32

Footnotes
1PSYCH 372 may be taken in place of NEUROSCI 301 with department permission.
2Professional or Technical Writing is recommended (e.g. ENGLISH 201 or 402)
3Computational Neuroscience Electives (the Hardware emphasis requires a minimum of 5 credits): At least 3 credits must be 300-400-level E E or CPT S courses. Approved courses include BIOLOGY 315, 321, 340, 438, 456; BIO ENG 481; CPT S 322, 434, 440, 443; E E 311, 321, 324, 341, 451, 464; MBIOS 303, 305, 401, 404, 413, 478; NEUROSCI 305, 333, 409; Psych 470, 490, 491; PHYSICS 466. Courses may not be used to fulfill more than one requirement. Other courses may be allowed by department consent. Please consult your advisor.

Computational Software Option (120 Credits)

Students may be admitted to the Neuroscience - Computational Software Option upon making their intention known to the department and enrollment in or credit for MATH 106 (or higher). To maintain admission to the major, students who have completed a minimum of 30 credits at WSU must maintain a 3.0 minimum GPA overall, and a minimum 3.0 cumulative GPA in BIOLOGY 107, CHEM 105, CHEM 106 or 116, MATH 171, MATH 172, NEUROSCI 301, NEUROSCI 302, and PHYSICS 201/211 or 205. Students must receive a grade of C or better in these courses and may repeat a maximum of three of these courses to maintain admission to the major.
First Year
First TermCredits
Arts [ARTS]3
CHEM 105 [PSCI]4
ENGLISH 101 [WRTG]3
HISTORY 105 [ROOT]3
MATH 171 [QUAN]4
Second TermCredits
BIOLOGY 107 [BSCI]4
CHEM 1064
CPT S 1214
MATH 1724
Second Year
First TermCredits
BIOLOGY 106 4
NEUROSCI 30113
PHYSICS 201/211 or 2054
PSYCH 105 [SSCI]3
Second TermCredits
CHEM 3454
CPT S 1224
NEUROSCI 302 3
PHYSICS 202/212 or 2064
Complete Writing Portfolio
Third Year
First TermCredits
Communication [COMM] or Written Communication [WRTG]23
E E 214 4
Humanities [HUM]3
MATH 2163
MBIOS 3014
Second TermCredits
CPT S 2233
Diversity [DIVR]3
NEUROSCI 4044
PSYCH 4903
Fourth Year
First TermCredits
CPT S 2242
CPT S 4403
NEUROSCI 4253
NEUROSCI 4261
NEUROSCI 430 [M]4
Second TermCredits
CPT S 3223
NEUROSCI 403 [M] 3
NEUROSCI 490 [CAPS]3
Computational Neuroscience Electives36

Footnotes
1PSYCH 372 may be taken in place of NEUROSCI 301 with department permission.
2Professional or Technical Writing is recommended (e.g. ENGLISH 201 or 402)
3Computational Neuroscience Electives (the Software emphasis requires a minimum of 6 credits): At least 3 credits must be 300-400-level E E or CPT S courses. Approved courses include BIOLOGY 315, 321, 340, 438, 456; BIO ENG 481; CPT S 322, 434, 440, 443; E E 311, 321, 324, 341, 451, 464; MBIOS 303, 305, 401, 404, 413, 478; NEUROSCI 305, 333, 409; Psych 470, 490, 491; PHYSICS 466. Courses may not be used to fulfill more than one requirement. Other courses may be allowed by department consent. Please consult your advisor.

General Option (120 Credits)

Students may be admitted to the Neuroscience - General Option upon making their intention known to the department and enrollment or credit for MATH 106 (or higher). To maintain admission to the major, students who have completed a minimum of 30 credits at WSU must maintain a 3.0 minimum GPA overall, and a 3.0 minimum cumulative GPA in BIOLOGY 107, CHEM 105, CHEM 106 or 116, MATH 140 or 171, NEUROSCI 301, NEUROSCI 302, PHYSICS 101/111 or 201/211 or 205, and PHYSICS 102/112 or 202/212 or 206 or CHEM 345. Students must receive a grade of C or better in these courses and may repeat a maximum of three of these courses to maintain admission to the major.
First Year
First TermCredits
CHEM 105 [PSCI]4
ENGLISH 101 [WRTG]3
MATH 140 [QUAN] or 171 [QUAN]4
PSYCH 105 [SSCI]3
Second TermCredits
BIOLOGY 107 [BSCI]4
CHEM 1064
Communication [COMM] or Written Communication [WRTG]13
HISTORY 105 [ROOT]3
Second Year
First TermCredits
BIOLOGY 106 4
CHEM 3454
NEUROSCI 30123
PHYSICS 101 or 2013
PHYSICS 111 or 2111
Second TermCredits
Arts [ARTS]3
Diversity [DIVR]3
NEUROSCI 3023
PHYSICS 102 or 2023
PHYSICS 112 or 2121
Neuroscience Electives34
Complete Writing Portfolio
Third Year
First TermCredits
Behavior Course43 or 4
BIOLOGY/MBIOS 3014
Statistics Course53 or 4
Neuroscience Electives35
Second TermCredits
Humanities [HUM]3
NEUROSCI 4044
Electives 68
Fourth Year
First TermCredits
NEUROSCI 430 [M]4
Electives 611
Second TermCredits
NEUROSCI 403 [M]3
NEUROSCI 490 [CAPS]3
Electives 69

Footnotes
1NEUROSCI/MBIOS 201 is recommended.
2PSYCH 372 may be taken in place of NEUROSCI 301 with department permission.
3Neuroscience Electives (9 credits): Approved courses include BIOLOGY 307, 315, 321, 333, 340, 352, 353, 354, 438, 456; MATH 340; MBIOS 303, 304, 305, 401, 404, 413; NEUROSCI 305, 333, 409, 425, 426; PHYSICS 466; PSYCH 265, 312, 333, 350, 361, 372, 384, 464, 470, 490, 491; VET PH 308. Other courses may be allowed by department consent. Courses may not be used to fulfill more than one requirement. Please see your advisor.
4Behavior Course: Choose one course from NEUROSCI 305, 333, or 409. Other courses may be allowed by department consent.
5Statistics Course: Choose one course from PSYCH 311, STAT 212, 360, 370, or 412.
6Elective choices should include 300-400 level coursework to meet the University minimum requirement of 40 upper division credits.

Honors Accelerated Pre-Veterinary Option (120 Credits)

This option has been established for admission of highly academically qualified students to the Doctor of Veterinary Medicine (D.V.M.) program at the Washington State University College of Veterinary Medicine (CVM). The program of study consists of three years of undergraduate coursework that fulfills the pre-veterinary neuroscience requirements followed by the four-year D.V.M. program. Satisfactory completion of this 7-year curriculum leads to the Bachelor of Science (B.S.) in Neuroscience and Doctor of Veterinary Medicine (D.V.M.).

All students who qualify for admission to the WSU Honors College are eligible to apply for pre-admission to the College of Veterinary Medicine after completion of one year in the pre-veterinary neuroscience curriculum. Interested applicants should identify themselves to the Honors College as soon as they decide to enroll at the University because the number of available seats in the B.S./D.V.M. program is limited. Early admission to the D.V.M. program requires approval of the CVM Admissions Committee. Accepted students are pre-admitted directly to the D.V.M. program. To maintain pre-admission into the D.V.M. program, accepted students must achieve an overall grade point average of 3.5 or better in all undergraduate coursework.

Students may be admitted to the Neuroscience Honors Accelerated Pre-Veterinary Option upon making their intention known to the department and enrollment in, or credit for, MATH 106 (or higher). To maintain admission to the major, students who have completed a minimum of 30 credits at WSU must maintain a 3.0 minimum GPA overall, and a 3.0 minimum cumulative GPA in each of the following courses: BIOLOGY 107; CHEM 105; CHEM 106 or 116; MATH 140 or 171; NEUROSCI 301, NEUROSCI 302; PHYSICS 101/111, 201/211 or 205; and PHYSICS 102/112, 202/212, 206, or CHEM 345. Students must receive a grade of C or better in these courses and may repeat a maximum of three of these courses to maintain admission to the major.

Completion of the degree requires completion of the HONORS Curriculum; a minimum of 90 undergraduate credits, including 30 upper-division credits; and one year of DVM coursework.
First Year
First TermCredits
BIOLOGY 1074
CHEM 1054
MATH 140 or 1714
Foreign Language (if needed)10-4
Second TermCredits
CHEM 106 or 11624
ENGLISH 2983
HONORS 27033
PHYSICS 1013
PHYSICS 1111
Foreign Language (if needed)10-4
Second Year
First TermCredits
BIOLOGY 1064
HONORS 2803
HONORS 29023
NEUROSCI 30143
PSYCH 10533
Second TermCredits
CHEM 3454
NEUROSCI 3023
PHYSICS 1023
PHYSICS 1121
Behavior Requirement53 or 4
Complete Writing Portfolio
Third Year
First TermCredits
BIOLOGY/MBIOS 3014
HONORS 3703
HONORS 39860 or 1
MBIOS 3034
NEUROSCI 430 [M]4
NEUROSCI 45072
Second TermCredits
HONORS 3803
HONORS 3903
HONORS 45071
NEUROSCI 403 [M]3
NEUROSCI 490 [CAPS]3
Statistics Course84
Fourth Year
First TermCredits
VET MED 5115
Additional D.V.M. Coursework910
Second TermCredits
VET MED 5205
VET MED 521103
Additional D.V.M. Coursework97

Footnotes
1Students must meet the Honors College Foreign Language requirement.
2Students who complete CHEM 116 fulfill the Honors College HONORS 290 requirement and another 3-credit course can be substituted.
3If HONORS 270 is taught by a member of the Psychology faculty, students may waive PSYCH 105. If PSYCH 105 is waived, students may need to take additional coursework to meet the 90-credit undergraduate minimum.
4PSYCH 372 may be taken in place of NEUROSCI 301 with department permission.
5Behavior Course: Choose one course from NEUROSCI 305, 333, or 409. Other courses may be allowed by department consent.
6HONORS 398 is an optional thesis-preparation course.
7NEUROSCI 450 is applied to the 3 credit HONORS 450 requirement.
8Statistics Course: Choose one course from PSYCH 311, STAT 212, 360, 370, or 412.
9Additional D.V.M. courses required in the first year of the D.V.M. program to satisfy the Neuroscience elective requirement for the B.S. in Neuroscience.
10VET MED 521 satisfies the Neuroanatomy (NEUROSCI 404) requirement for the B.S. in Neuroscience.

Pre-Medical and Pre-Dental Option (120 Credits)

Students may be admitted to the Neuroscience - Pre-Medical and Pre-Dental Option upon making their intention known to the department and enrollment in or credit for MATH 106 (or higher). To maintain admission to the major, students who have completed a minimum of 30 credits at WSU must maintain a 3.0 minimum GPA overall, and a 3.0 minimum cumulative GPA in BIOLOGY 107, CHEM 105, CHEM 106 or 116, MATH 140 or 171, NEUROSCI 301, NEUROSCI 302, PHYSICS 101/111 or 201/211 or 205, and PHYSICS 102/112 or 202/212 or 206 or CHEM 345. Students must receive a grade of C or better in these courses and may repeat a maximum of three of these courses to maintain admission to the major.

Students are encouraged to take the MCAT after completion of the third year.
First Year
First TermCredits
CHEM 105 [PSCI]4
ENGLISH 101 [WRTG]3
MATH 140 [QUAN] or 171 [QUAN]4
PSYCH 105 [SSCI]3
Second TermCredits
BIOLOGY 107 [BSCI]4
CHEM 1064
Communication [COMM] or Written Communication [WRTG]13
HISTORY 105 [ROOT]3
Second Year
First TermCredits
BIOLOGY 1064
CHEM 3454
NEUROSCI 30123
PHYSICS 101 or 2013
PHYSICS 111 or 2111
Second TermCredits
Arts [ARTS]3
CHEM 3484
Diversity [DIVR]3
NEUROSCI 3023
PHYSICS 102 or 2023
PHYSICS 112 or 2121
Complete Writing Portfolio
Third Year
First TermCredits
Behavior Course33 or 4
MBIOS 3034
Statistics Course43 or 4
Neuroscience Electives55
Second TermCredits
BIOLOGY/MBIOS 3014
Humanities [HUM]3
NEUROSCI 4044
Electives64
Fourth Year
First TermCredits
NEUROSCI 430 [M]4
Electives611
Second TermCredits
NEUROSCI 403 [M]3
NEUROSCI 490 [CAPS]3
Electives69

Footnotes
1NEUROSCI/MBIOS 201 is recommended.
2PSYCH 372 may be taken in place of NEUROSCI 301 with department permission.
3Behavior Course: Choose one course from NEUROSCI 305, 333, or 409. Other courses may be allowed by department consent.
4Statistics Course: Choose one course from PSYCH 311, STAT 212, 360, 370, or 412.
5Neuroscience Electives (5 credits): Approved courses include BIOLOGY 307, 315, 321, 333, 340, 352, 353, 354, 438, 456; MATH 340; MBIOS 304, 305, 401, 404, 413; NEUROSCI 305, 333, 409, 425, 426; PHYSICS 466; PSYCH 265, 312, 333, 350, 361, 372, 384, 464, 470, 490, 491; VET PH 308. Courses may not be used to fulfill more than one requirement. Other courses may be allowed by department consent. Please see your advisor.
6Electives: Consult your advisor regarding elective courses that may be required or recommended for admission into your future health-professions program.

Pre-Veterinary Option (120 Credits)

Students may be admitted to the Neuroscience - Pre-Veterinary Option upon making their intention known to the department and enrollment in or credit for MATH 106 (or higher). To maintain admission to the major, students who have completed a minimum of 30 credits at WSU must maintain 3.0 minimum GPA overall, and a 3.0 minimum cumulative GPA in BIOLOGY 107, CHEM 105, CHEM 106 or 116, MATH 140 or 171, NEUROSCI 301, NEUROSCI 302, PHYSICS 101/111 or 201/211 or 205, and PHYSICS 102/112 or 202/212 or 206 or CHEM 345. Students must receive a grade of C or better in these courses and may repeat a maximum of three of these courses to maintain admission to the major.

Students are encouraged to take the GRE after completion of the third year and apply to Veterinary School by the end of the first term of the fourth year.
First Year
First TermCredits
CHEM 105 [PSCI]4
ENGLISH 101 [WRTG]3
MATH 140 [QUAN] or 171 [QUAN]4
PSYCH 105 [SSCI]3
Second TermCredits
BIOLOGY 107 [BSCI]4
CHEM 1064
Communication [COMM] or Written Communication [WRTG]13
HISTORY 105 [ROOT]3
Second Year
First TermCredits
BIOLOGY 106 4
CHEM 3454
NEUROSCI 30123
PHYSICS 101 or 2013
PHYSICS 111 or 2111
Second TermCredits
BIOLOGY/MBIOS 3014
Diversity [DIVR]3
NEUROSCI 3023
PHYSICS 102 or 2023
PHYSICS 112 or 2121
Complete Writing Portfolio
Third Year
First TermCredits
Behavior Course33 or 4
MBIOS 3034
Statistics Course43 or 4
Neuroscience Electives55
Second TermCredits
Arts [ARTS]3
Humanities [HUM]3
NEUROSCI 4044
Electives67
Fourth Year
First TermCredits
NEUROSCI 430 [M]4
Electives612
Second TermCredits
NEUROSCI 403 [M]3
NEUROSCI 490 [CAPS]3
Electives69

Footnotes
1NEUROSCI/MBIOS 201 is recommended.
2PSYCH 372 may be taken in place of NEUROSCI 301 with department permission.
3Behavior Course: Choose one course from NEUROSCI 305, 333, or 409. Other courses may be allowed by department consent.
4Statistics Course: Choose one course from PSYCH 311, STAT 212, 360, 370, or 412.
5Neuroscience Electives (5 credits): Approved courses include BIOLOGY 307, 315, 321, 333, 340, 352, 353, 354, 438, 456; MATH 340; MBIOS 301, 304, 305, 401, 404, 413; NEUROSCI 305, 333, 409, 425, 426; PHYSICS 466; PSYCH 265, 312, 333, 350, 361, 372, 384, 464, 470, 490, 491; VET PH 308. Other courses may be allowed by department consent. Courses may not be used to fulfill more than one requirement. Please consult your advisor.
6Elective choices should include 300-400 level coursework to meet the University minimum requirement of 40 upper division credits. Consult your advisor regarding elective courses that may be required or recommended for admission to a DVM program.


Minors

Neuroscience

Students may apply for the minor in Neuroscience once they have completed 60 credits and have a 2.75 GPA. However, they may take minor coursework at any time as long as they meet the prerequisites. The minor in Neuroscience requires 16 credits with at least 13 credits at or above the 300-level. Courses needed to satisfy the minor must include NEUROSCI 301; three credits selected from NEUROSCI 305, 333, or  409; at least six credits selected from the following: NEUROSCI 403, 404, and 430; and up to four credits of neuroscience related elective coursework. Approved Neuroscience electives include: BIOLOGY 301, 315, 321, 340, 352, 353, 354, 438, 456; MATH 340; MBIOS 301, 304, 305, 401, 404, 413; NEUROSCI 305, 409, 425, 426; PSYCH 265, 312, 333, 350, 361, 372, 384, 464, 470, 490, 491; PHYSICS 466; VET PH 308. Upon the approval of the student's advisor, a student with a minor in neuroscience may include 500-level courses in the minor program, provided the student meets the graduate study requirements and, prior to registration, obtains the consent of the faculty Neuroscience minor.  Credits for the minor must include a minimum of  9 credits of upper-division work taken in residence at WSU or through WSU-approved education abroad or educational exchange courses.



Certificates

Certificate in the Business of Biomedicine

The certificate in the Business of Biomedicine prepares students for careers in the dynamic, interdisciplinary world of biomedical business. To complete the certificate, students must complete a minimum of 16 credits consisting of two core courses and three additional electives with a GPA of 2.5 or better. Required core courses are BIOLOGY/MBIOS 301 and MGMT 301. Fulfillment of elective emphasis courses requires completion of one of ACCTG 220, ACCTG 230, FIN 223, or ENTRP 426; AND one of ENTRP 490, COMSTRAT 380, or COMSTRAT 395; AND one of NEUROSCI 302 or MBIOS 303 or another course as approved by your advisor.


Courses

The online catalog includes the most recent changes to courses and degree requirements that have been approved by the Faculty Senate, including changes that are not yet effective. Courses showing two entries of the same number indicate that the course information is changing. The most recently approved version is shown first, followed by the older version, in gray, with its last-effective term preceding the course title. Courses shown in gray with only one entry of the course number are being discontinued. Course offerings by term can be accessed by clicking on the term links when viewing a specific campus catalog.


Neuroscience (NEUROSCI)

(Select Campus to see schedule links)


105 [BSCI] Meet Your Brain 3 Introduction to the brain and nervous system and how they are affected by your environment and everyday activities.

138 Neuroscience Seminar 1 May be repeated for credit; cumulative maximum 2 credits. Introduces new students to individual faculty research interests and helps students link personal interests to academic majors. S, F grading.

150 [BSCI] Art and the Brain 3 Explore networks involved in sensory perception; learn how art integrates perceptual experiences into works that affect our understanding of ourselves.

201 [COMM] Introduction to Communication in the Molecular Life Sciences 3 (1-4) Course Prerequisite: BIOLOGY 106 with a C or better or BIOLOGY 107 with a C or better. Analysis of primary literature and an introduction to scientific communication skills in the molecular life sciences. (Crosslisted course offered as NEUROSCI 201, MBIOS 201). Recommended preparation: Pre-admitted or admitted major in Biochemistry, Genetics & Cell Biology, Microbiology, or Neuroscience.

275 Special Topics: Study Abroad V 1-15 May be repeated for credit. S, F grading.

301 Foundations of Neuroscience I 3 Course Prerequisite: BIOLOGY 107, CHEM 105, or NEUROSCI 138. Enrollment in NEUROSCI 301 not allowed if credit already earned for PSYCH 372. Structure and function of the nervous system from single neurons to behavior.

302 Foundations of Neuroscience II 3 Course Prerequisite: CHEM 106; BIOLOGY 107; NEUROSCI 301 or PSYCH 372. In-depth concepts, analysis, and discussion of the experimental foundations for understanding nervous system function.

305 Neurons, Genes, and Behavior 3 Course Prerequisite: NEUROSCI 301, 302, MBIOS 301, or PSYCH 372. In-depth exploration of the organization of the brain, and how this organization underlies behavior.

333 Techniques and Experimental Design in Neuroscience Research 4 (3-3) Course Prerequisite: NEUROSCI 301 or PSYCH 372; MATH 140, 171, PSYCH 311, STAT 212, 360, 370, or 412. Representative modern neuroscience experiments are explored from a conceptual, technical, and design perspective.

403 [M] Cellular Neurobiology 3 Course Prerequisite: NEUROSCI 302; MBIOS 301; admitted to the major or minor in Neuroscience. Cellular and molecular interactions occurring within the nervous system. Recommended preparation: NEUROSCI 430.

404 Neuroanatomy 4 (3-3) Course Prerequisite: NEUROSCI 302. Fundamental principles of the organization and plans of circuitry of the nervous system.

409 Affective Neuroscience 3 Course Prerequisite: NEUROSCI 302 or PSYCH 372. Brain mechanisms of human and animal emotions. Credit not granted for both NEUROSCI 409 and NEUROSCI 509. Offered at 400 and 500 level.

425 Foundations of Medical Physiology 3 Course Prerequisite: BIO ENG 210, MBIOS 301, MBIOS 303, NEUROSCI 301, NEUROSCI 302, or PSYCH 372. Integrated functioning of organ systems, focusing on mechanisms of disease, organ dysfunction, and disturbances to whole-animal homeostasis and health. (Crosslisted course offered as NEUROSCI 425, VET PH 425).

426 Foundations of Medical Physiology Lab 1 (0-3) Course Prerequisite: Concurrent enrollment in NEUROSCI 425. Optional laboratory component of NEUROSCI/VET PH 425. Practical analysis of organ function and health using medical diagnostic equipment and clinical cases. (Crosslisted course offered as NEUROSCI 426,VET PH 426).

430 [M] Principles of Neurophysiology 4 (3-3) Course Prerequisite: NEUROSCI 302; 4 credits of PHYSICS 102 or 202, or PHYSICS 102 and 112, or PHYSICS 202 and 212, or PHYSICS 206. Advanced exploration of the principles underlying cellular, sensory, motor and integrative functions of the nervous system. Recommended preparation: MBIOS 303.

450 Honors Thesis Research V 1-3 Course Prerequisite: Admitted to the major in Neuroscience; admitted to Honors College. Laboratory research with emphasis on honors thesis or project directed by faculty.

480 Special Topics: Study Abroad V 1-15 May be repeated for credit. S, F grading.

490 [CAPS] Senior Project 3 Course Prerequisite: Admitted to the major in Neuroscience; senior standing. Senior capstone course for students majoring in Neuroscience.

491 Senior Project-Poster 1 Course Prerequisite: Admitted to the major in Neuroscience; senior standing. Research project poster or oral presentation. S, F grading.

495 Directed Research V 1-3 May be repeated for credit. Course Prerequisite: Admitted to the major or minor in Neuroscience. Introduction to neuroscience research literature.

496 Special Topics V 1-3 May be repeated for credit; cumulative maximum 6 credits. Course Prerequisite: NEUROSCI 302 with a C or better or junior standing. Reading, discussion, and exploration of contemporary topics in Neuroscience.

499 Special Problems V 1-3 May be repeated for credit; cumulative maximum 6 credits. Course Prerequisite: Admitted to the major or minor in Neuroscience. Independent study conducted under the jurisdiction of an approving faculty member; may include independent research studies in technical or specialized problems; selection and analysis of specified readings; development of a creative project; or field experiences. S, F grading.

509 Affective Neuroscience 3 Brain mechanisms of human and animal emotions. Credit not granted for both NEUROSCI 409 and NEUROSCI 509. Offered at 400 and 500 level.

520 Fundamentals of Neuroscience 4 (3-3) Functional aspects of the brain from cell membrane to higher integrative processes. Cooperative: Open to UI degree-seeking students.

529 Integrative Neuroscience 3 Basic biochemical processes in the nervous system and their significance for normal and abnormal function.

531 Neuroscience Laboratory Rotation 1 (0-3) May be repeated for credit; cumulative maximum 2 credits. Fourteen-week rotation through each of two research laboratories; learning procedures and techniques in neuroscience. S, F grading.

540 Special Topics in Integrative Neuroscience V 1-3 May be repeated for credit; cumulative maximum 6 credits. Concepts and controversies in neuroscience involving integrative properties of cell systems. Cooperative: Open to UI degree-seeking students.

541 Special Topics in Cellular and Molecular Neuroscience V 1-3 May be repeated for credit; cumulative maximum 6 credits. Concepts and controversies in neuroscience that involve nerve cell function and regulation. Cooperative: Open to UI degree-seeking students.

542 Special Topics in Interdisciplinary Neuroscience V 1-3 May be repeated for credit; cumulative maximum 6 credits. Concepts and controversies in neuroscience that revolve around traditional approaches to nervous system study. Cooperative: Open to UI degree-seeking students.

543 Special Topics in Behavioral/Clinical Neuroscience V 1-3 May be repeated for credit; cumulative maximum 6 credits. Concepts and controversies in neuroscience that involve normal and pathological aspects of behavior. Cooperative: Open to UI degree-seeking students.

563 Deconstruction of Research 3 Course Prerequisite: Graduate standing in a WSU biomedical based graduate program. Nature and development of scientific investigation through oral and written avenues, and methods of critical analyses applied to questions of biomedical interest. (Crosslisted course offered as NEUROSCI 563, GLANHLTH 563, MBIOS 563, VET MICR 563, VET PATH 563, VET PH 563).

564 Topics in Biomedical Experimentation V 1-3 May be repeated for credit; cumulative maximum 6 credits. Examination of the philosophy of experimental design and practical application and analysis of various experimental approaches in biomedical research. Recommended preparation: graduate standing in a WSU biomedical-based program, and an advanced undergraduate or graduate statistics course. (Crosslisted course offered as NEUROSCI 564, GLANHLTH 564, MBIOS 564, PHIL 564, VET MICR 564, VET PATH 564, VET PH 564).

590 Seminar 1 May be repeated for credit; cumulative maximum 7 credits. Presented by advanced graduate students and faculty (both in INP and around WSU) on their research areas. (Crosslisted course offered as NEUROSCI 590, VET PH 590.) S, F grading.

592 Research Writing and Seminar 2 May be repeated for credit; cumulative maximum 6 credits. Course Prerequisite: Graduate student in Neuroscience program. Essentials of oral and written scientific communication.

600 Special Projects or Independent Study V 1-18 May be repeated for credit. Independent study, special projects, and/or internships. Students must have graduate degree-seeking status and should check with their major advisor before enrolling in 600 credit, which cannot be used toward the core graded credits required for a graduate degree. S, F grading.

700 Master's Research, Thesis, and/or Examination V 1-18 May be repeated for credit. Independent research and advanced study for students working on their master's research, thesis and/or final examination. Students must have graduate degree-seeking status and should check with their major advisor/committee chair before enrolling for 700 credit. S, U grading.

800 Doctoral Research, Dissertation, and/or Examination V 1-18 May be repeated for credit. Course Prerequisite: Admitted to the Neuroscience PhD program. Independent research and advanced study for students working on their doctoral research, dissertation and/or final examination. Students must have graduate degree-seeking status and should check with their major advisor/committee chair before enrolling for 800 credit. S, U grading.

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